Evolutionary Anachronisms in The Western Palearctic – Part I: Puzzling Pomes

In their 1982 paper Neotropical Anachronisms: The Fruits the Gomphotheres Ate, Daniel H. Janzen and Paul S. Martin defined it as a trait of a plant that is inexplicable unless seen in the backdrop of its evolutionary past. For instance, Gymnocladus dioicus, a tree in the legume plant family Fabaceae that is native to the eastern United States, produces seed pods that are poisonous to mammals, unbreakable to rodents and impervious to water, yet depend on all of these for dispersal since the seeds, the largest in the continental United States, are too heavy to be carried by wind anywhere far. As a result, each year the parent tree will produce pods that fall to the ground, where they slowly decompose over the years, even in seemingly natural habitat. This is odd because the fruit of any plant is always intended as a diaspore. Plants have outbid each other over millions of years in attempts to produce the most sophisticated designs that will allow their unborn offspring to travel and germinate a preferably long distance away from the parent. So, if the Kentucky coffeetree, as it is also called, fails so miserably at dispersing seeds away from the parent tree, despite an elaborate diaspore, one is compelled to ask, why?.